The American Canine Association ( ACA ) Search and Rescue Teams of 911. Actual pictures
taken by American Canine Association ( ACA ) Search and Rescue Team members during
the ACA's vital and heroic rescue efforts at Ground Zero in New York, NY. American
Canine Association ( ACA ) Search and Rescue Team Leader Bob Yarnall, Jr. is also
the President of American Canine Association ( ACA ). American Canine Association
( ACA ) Search and Rescue Team is a non-
Domestic dogs often display the remnants of countershading, a common natural camouflage pattern. The general theory of countershading is that an animal that is lit from above will appear lighter on its upper half and darker on its lower half, where it will usually be in its own shade. This is a pattern that prey can learn to watch for. A countershaded animal will have dark coloring on its upper surfaces and light coloring below, which reduces its general visibility. Thus many breeds will have an occasional "blaze", stripe, or "star" of white fur on their chest or underside.
Tail
There are many different shapes for dog tails: straight, straight up, sickle, curled,
or cork-
Types and breeds
Main article: Dog breeds
Further information: Dog type
While all dogs are genetically very similar, natural selection and selective breeding have reinforced certain characteristics in certain populations of dogs, giving rise to dog types and dog breeds. Dog types are broad categories based on function, genetics, or characteristics.
Dog breeds are groups of animals that possess a set of
inherited characteristics that distinguishes them from other animals within the
same species. Modern dog breeds are non-
Health
Main article: Dog health
Further information: Category:Dog health
See also: Canine Vector-
Dogs are susceptible to various diseases, ailments, and poisons, some of which can affect humans. To defend against many common diseases, dogs are often vaccinated.
Some breeds of dogs are prone to certain genetic ailments
such as elbow or hip dysplasia, blindness, deafness, pulmonic stenosis, cleft palate, and trick
knees. Two serious medical
conditions particularly affecting dogs are pyometra, affecting unspayed females of all types and ages, and bloat, which affects
the larger breeds or deep-
Dogs are also vulnerable to some of the same health conditions as humans, including diabetes, dental and heart disease, epilepsy, cancer, hypothyroidism, and arthritis.
Mortality
Main article: Aging in dogs
The typical lifespan of dogs varies widely among breeds, but for most the median longevity, the age at which half the dogs in a population have died and half are still alive, ranges from 10 to 13 years. Individual dogs may live well beyond the median of their breed.
The breed with the shortest lifespan (among breeds for
which there is a questionnaire survey with a reasonable sample size) is the
Dogue
de Bordeaux, with a median
longevity of about 5.2 years, but several breeds, including Miniature Bull Terriers, Bloodhounds, and Irish
Wolfhounds are nearly as
short-
The longest-
Predation
Although wild dogs, like wolves, are apex predators, they can be killed in territory disputes with wild animals. Furthermore, in areas where both dogs and other large predators live, dogs can be a major food source for big cats or canines. Reports from Croatia indicate that dogs are killed more frequently than sheep. Wolves in Russia apparently limit feral dog populations. In Wisconsin, more compensation has been paid for dog losses than livestock. Some wolf pairs have been reported to prey on dogs by having one wolf lure the dog out into heavy brush where the second animal waits in ambush. In some instances, wolves have displayed an uncharacteristic fearlessness of humans and buildings when attacking dogs, to the extent that they have to be beaten off or killed. Coyotes and big cats have also been known to attack dogs. Leopards in particular are known to have a predilection for dogs, and have been recorded to kill and consume them regardless of the dog's size or ferocity. Tigers in Manchuria, Indochina, Indonesia, and Malaysia, are reputed to kill dogs with the same vigor as leopards. Striped Hyenas are major predators of village dogs in Turkmenistan, India, and the Caucasus.
The eyes of different breeds of dogs have different shapes,
dimensions, and retina configurations.[ Many
long-
Hearing
The frequency range of dog hearing is approximately 40 Hz to 60,000 Hz,[ which means that dogs can detect sounds far beyond the upper limit of the human auditory spectrum. Additionally, dogs have ear mobility which allows them to rapidly pinpoint the exact location of a sound. Eighteen or more muscles can tilt, rotate, raise, or lower a dog's ear. A dog can identify a sound's location much faster than a human can, as well as hear sounds at four times the distance.
Smell
While the human brain is dominated by a large visual
cortex, the dog brain is
dominated by an olfactory cortex. The olfactory bulb in dogs is roughly forty times bigger than the olfactory
bulb in humans, relative to total brain size, with 125 to 220 million smell-
Physical characteristics
Coat
Main article: Coat (dog)
The coats of domestic dogs are either "double", made up of a coarse topcoat and a soft undercoat, like a wolf, or "single", with the topcoat only. Dogs with double coats tend to originate in colder climates.
The incidence of dog bites, and especially fatal dog bites, is extremely rare in America considering the number of pet dogs in the country. Fatalities from dog bites occur in America at the rate of one per four million dogs. A Colorado study found that bites in children were less severe than bites in adults. The incidence of dog bites in the US is 12.9 per 10,000 inhabitants, but for boys aged 5 to 9 the incidence rate is 60.7 per 10,000. Moreover, children have a much higher chance to be bitten in the face or neck. Sharp claws with powerful muscles behind them can lacerate flesh in a scratch that can lead to serious infections.
In the UK between 2003 and 2004, there were 5,868 dog attacks on humans resulting in 5,770 working days lost in sick leave.
Biology
Main article: Dog anatomy
Domestic dogs have been selectively bred for millennia for
various behaviors, sensory capabilities, and physical attributes. Modern
dog
breeds show more variation
in size, appearance, and behavior than any other domestic animal. Nevertheless,
their morphology is based on that of their wild ancestors, gray wolves. Dogs are
predators and scavengers, and like many other predatory mammals, the dog has
powerful muscles, fused wrist bones, a cardiovascular system that supports both
sprinting and endurance, and teeth for catching and tearing. Dogs are highly
variable in height and weight. The smallest known adult dog was a Yorkshire Terrier, that stood only 6.3 cm (2.5 in) at the shoulder,
9.5 cm (3.75 in) in length along the head-
Senses
Sight
Like most mammals, dogs are dichromats and have color vision equivalent to red-
The dog's visual system has evolved to aid proficient hunting. While a dog's visual acuity is poor (that of a poodle's has been estimated to translate to a Snellen rating of 20/75), their visual discrimination for moving objects is very high; dogs have been shown to be able to discriminate between humans (e;g. identifying their owner) from distances up to a mile.[47] As crepuscular hunters, dogs often rely on their vision in low light situations: they have very large pupils, a high density of rods in the fovea, an increased flicker rate, and a tapetum lucidum.[47] The tapetum is a reflective surface behind the retina that reflects light back to give the photoreceptors a second chance to catch the photons.
Sports and shows
Owners of dogs often enter them in competitions such as breed conformation shows or sports, including racing and sledding.
In conformation shows, also referred to as breed shows, a judge familiar with the specific dog breed evaluates individual purebred dogs for conformity with their established breed type as described in the breed standard. As the breed standard only deals with the externally observable qualities of the dog (such as appearance, movement, and temperament), separately tested qualities (such as ability or health) are not part of the judging in conformation shows.
As a food source
See also: Dog meat
Dog meat is consumed in some East
Asian countries, including
China, Korea, and Vietnam, a practice that dates back to antiquity. In Korea, the
primary dog breed raised for meat, the nureongi,
differs from those breeds raised for pets which Koreans may keep in their homes.
The most popular Korean dog dish is gaejang-
Other cultures, such as Polynesia and Pre-
Health risks to humans
In the USA, dogs cause 88% of the 86,000 falls caused by pets.[citation needed]
Dog feces can cause a number of human diseases, including toxocariasis, which can cause blindness, and can also cause human hookworm. In the United States, about 10,000 cases of Toxocara infection are reported in humans each year. Almost 14% of the US population is infected with Toxocara, a parasite of dogs and cats that can be passed from animals to humans.
The dog quickly became ubiquitous across culture in all parts of the world, and was extremely valuable to early human settlements. For instance, it is believed that the successful emigration across the Bering Strait might not have been possible without sled dogs. Dogs perform many roles for people, such as hunting, herding, protection, and, more recently, assisting handicapped individuals. Currently, there are estimated to be 400 million dogs in the world.
Over the 15,000 year span that the dog had been
domesticated, it diverged into only a handful of landraces, groups of similar animals whose morphology and behavior have been shaped by environmental factors and functional roles. As the modern understanding of
genetics developed, humans began to intentionally breed dogs for a wide range of
specific traits. Through this process, the dog has developed into hundreds of
varied breeds, and shows more behavioral and morphological variation
than any other land mammal.[5] For
example, height measured to the withers ranges from a few inches in the Chihuahua to a few feet in the Irish
Wolfhound; color varies
from white through grays (usually called "blue'") to black, and browns from
light (tan) to dark ("red" or "chocolate") in a wide variation of patterns;
coats
can be short or long, coarse-
Etymology and related terminology
Dog is the common use term that refers to members of the subspecies Canis lupus familiaris. The term is sometimes used to refer to a wider range of species: it can be used to refer to any mammal belonging to the family Canidae, which includes wolves, foxes, jackals, and coyotes; it can be used to refer to the subfamily of Caninae, or the genus Canis, also often called the "true dogs". Some members of the family have "dog" in their common names, such as the Raccoon Dog and the African Wild Dog. A few animals have "dog" in their common names but are not canids, such as the prairie dog and the dog fish.
The English word dog comes
from Middle
English dogge, from Old
English docga, a "powerful dog breed". The term may derive from
Proto-
Dog
ousted traditional English hound
by the 16th century, before which time it had the meaning of "dog" rather than
modern "hunting dog", as in other Germanic languages – it is cognate to German Hund,
Dutch hond, common Scandinavian hund, and Icelandic hundur.
Hound itself is derived from the Proto-
In breeding circles, a male canine is referred to as a dog, while a female is called a bitch. A group of offspring is a litter. The father of a litter is called the sire, and the mother is called the dam. Offspring are generally called pups or puppies until they are about a year old. The process of birth is whelping.
Taxonomy
The domestic dog was originally classified as Canis familiaris and Canis familiarus domesticus by Carolus Linnaeus in 1758, and is currently classified as Canis lupus familiaris, a subspecies of the gray wolf Canis lupus, by the Smithsonian Institution and the American Society of Mammalogists. Overwhelming evidence from behavior, vocalizations, morphology, and molecular biology led to the contemporary scientific understanding that a single species, the gray wolf, is the common ancestor for all breeds of domestic dogs, however the timeframe and mechanisms by which dogs diverged are controversial.
History and evolution
Main article: Origin of the domestic dog
Domestic dogs inherited a complex social hierarchy and behaviors from their wolf ancestors. Dogs are pack animals with a complex set of behaviors related to determining each dog's position in the social hierarchy, and they exhibit various postures and other means of nonverbal communication that reveal their states of mind. These sophisticated forms of social cognition and communication may account for their trainability, playfulness, and ability to fit into human households and social situations, and these attributes have earned dogs a unique relationship with humans despite being potentially dangerous apex predators.
Although experts largely disagree over the details of dog domestication, it is agreed that human interaction played a significant role in shaping the subspecies. Shortly after domestication, dogs became ubiquitous in human populations, and spread throughout the world. Emigrants from Siberia likely crossed the Bering Strait with dogs in their company, and some experts suggest that use of sled dogs may have been critical to the success of the waves that entered North America roughly 12,000 years ago. Dogs were an important part of life for the Athabascan population in North America, and were their only domesticated animal. Dogs also carried much of the load in the migration of the Apache and Navajo tribes 1,400 years ago. Use of dogs as pack animals in these cultures often persisted after the introduction of the horse to North America.
The current consensus among biologists and archaeologists is that no one can be sure when dogs were domesticated. There is conclusive evidence that dogs genetically diverged from their wolf ancestors at least 15,000 years ago but some believe domestication to have occurred earlier. It is not known whether humans domesticated the gray wolf as such to initiate dog's divergence from its ancestors, or whether dog's evolutionary path took a different course already prior to domestication. Lately the latter view has gained proponents such as biologists Raymond and Lorna Coppinger. They theorize that some wolves started gathering around the campsites of the paleolithical man to scavenge his refuse. There, an evolutionary pressure developed that favored those who were less frightened by and keener in approaching humans.
The bulk of the scientific evidence for the evolution of the domestic dog stems from archaeological findings and mitochondrial DNA studies. The divergence date of roughly 15000 years ago is based in part on archaeological evidence that demonstrates that the domestication of dogs occurred more than 15,000 years ago, and some genetic evidence indicates that the domestication of dogs from their wolf ancestors began in the late Upper Paleolithic close to the Pleistocene/Holocene boundary, between 17,000 and 14,000 years ago. But there is a wide range of other, contradictory findings that make this issue controversial.
Archaeological evidence plays a large role in this debate. In 2008, a team of international scientists released findings from an excavation at Goyet Cave in Belgium declaring that a large, toothy canine existed 31,700 years ago and ate a diet of horse, musk ox and reindeer. Prior to this Belgium discovery, the earliest dog fossils were two large skulls from Russia and a mandible from Germany, that dated from roughly 14,000 years ago. Remains of smaller dogs from Natufian cave deposits in the Middle East have been dated to around 10,000 to 12,000 years ago. There is a great deal of archaeological evidence for dogs throughout Europe and Asia around this period and through the next two thousand years (roughly 8,000 to 10,000 years ago), with fossils uncovered in Germany, the French Alps, and Iraq, and cave paintings in Turkey.
Thus, the archaeological evidence suggests that the latest dogs could have diverged from wolves was roughly 15000 years ago, although it is possible that they diverged much earlier.
DNA studies have provided a wider range of possible
divergence dates, from 15,000 to 40,000 years ago,[20] to
as much as 100,000 to 140,000 years ago. This evidence depends on a number of
assumptions that may be violated. Genetic studies are based on comparisons of
genetic diversity between species, and depend on a calibration date. Many
estimates of divergence dates from DNA evidence use an estimated wolf-
Genetic analyses indicate all dogs are likely descended from a handful of domestication events with a small number of founding females, although there is evidence that domesticated dogs interbred with local populations of wild wolves on several occasions. Data suggests that dogs first diverged from wolves in East Asia, and that these domesticated dogs then quickly migrated throughout the world, reaching the North American continent around 8000 B.C. The oldest groups of dogs, which show the greatest genetic variability and are the most similar to their wolf ancestors, are primarily Asian and African breeds, including the Basenji, Lhasa Apso, and Siberian Husky. Some breeds that were thought to be very old, such as the Pharaoh Hound, Ibizan Hound, and Norwegian Elkhound, are now known to have been created more recently.
There is a great deal of controversy surrounding the evolutionary framework for the domestication of dogs.[3] Although it is widely claimed that "man domesticated the wolf," man may not have taken such a proactive role in the process. The nature of the interaction between man and wolf that led to domestication is unknown and controversial. At least three early species of the Homo genus began spreading out of Africa roughly 400,000 years ago, and thus lived for a considerable period in contact with canine species. Despite this, there is no evidence of any adaptation of canine species to the presence of the close relatives of modern man. If dogs were domesticated, as believed, roughly 15,000 years ago, the event (or events) would have coincided with a large expansion in human territory and the development of agriculture. This has led some biologists to suggest that one of the forces that led to the domestication of dogs was a shift in human lifestyle in the form of established human settlements. Permanent settlements would have coincided with a greater amount of disposable food and would have created a barrier between wild and anthropogenic canine populations.
Roles with humans
Early roles
This section requires expansion.
As pets
This section requires expansion.
Work
Dogs have lived and worked with humans in so many roles that they have earned the unique nickname, "man's best friend", a phrase which is used in other languages as well. They have been bred for herding livestock, hunting (e.g. pointers and hounds), rodent control, guarding, helping fishermen with nets, and pulling loads, in addition to their roles as companions.
Service dogs such as guide dogs, utility dogs, assistance dogs, hearing dogs, and psychological therapy dogs provide assistance to individuals with physical or mental disabilities. Some dogs owned by epileptics have been shown to alert their handler when the handler shows signs of an impending seizure, sometimes well in advance of onset, allowing the owner to seek safety, medication, or medical care.
Diet
See also: Dog food
Golden Retriever eating a pig's foot.
Despite its descent from wolves, the domestic dog is an
omnivore, though it is classified in the order Carnivora. Unlike an obligate carnivore, such as a member of the
cat
family with its shorter
small
intestine, a dog is
neither dependent on meat-
Reproduction
Main article: Canine reproduction
In domestic dogs, sexual maturity begins to happen around age six to twelve months for both males and females, although this can be delayed until up to two years old for some large breeds. This is the time at which female dogs will have their first estrous cycle. They will experience subsequent estrous cycles biannually, during which the body prepares for pregnancy. At the peak of the cycle, females will come into estrus, being mentally and physically receptive to copulation. Because the ova survive and are capable of being fertilized for a week after ovulation, it is possible for a female to mate with more than one male.
Dogs bear their litters roughly 56 to 72 days after fertilization, with an average of 63 days, although the length of gestation can vary. An average litter consists of about six puppies, though this number may vary widely based on the breed of dog. Toy dogs generally produce from one to four puppies in each litter, while much larger breeds may average as many as twelve.
Some dog breeds have acquired traits through selective breeding that interfere with reproduction. Male French Bulldogs, for instance, are incapable of mounting the female. For many dogs of this breed, the female must be artificially inseminated in order to reproduce.
Intelligence and behavior
Intelligence
Main article: Dog intelligence
The domestic dog has a predisposition to exhibit a social intelligence that is uncommon in the animal world. Dogs are capable of learning in a number of ways, such as through simple reinforcement (e.g. classical or operant conditioning) and by observation.
Dogs go through a series of stages of cognitive development. They are not born with the understanding that objects which are not being actively perceived still remain in existence, called object permanence. This occurs as the infant learns to interact intentionally with objects around it. For dogs, this occurs at roughly 8 weeks of age.
Puppies learn behaviors quickly by following examples set
by experienced dogs This form of intelligence is not peculiar to those tasks
dogs have been bred to perform, but can be generalized to myriad abstract
problems. For example, Dachshund puppies who watched an experienced dog pull a
cart by tugging on an attached piece of ribbon in order to get a reward from
inside the cart learned the task fifteen times faster than those who were left
to solve the problem on their own. Dogs can also learn by mimicking human
behaviors. In one study, puppies were presented with a box, and shown that when
a handler pressed a lever, a ball would roll out of the box. The handler then
allowed the puppy to play with the ball, making it an intrinsic reward. The pups
were then allowed to interact with the box. Roughly three-
Dogs also demonstrate sophisticated social
cognition by associating
behavioral cues with abstract meanings. One such class of social cognition
involves the understanding that others are conscious agents. Research has shown
that dogs are capable of interpreting subtle social cues, and appear to
recognize when a human or dog's attention is focused on them. To test this,
researchers devised a task in which a reward was hidden underneath one of two
buckets. The experimenter then attempted to communicate with the dog to indicate
the location of the reward by using a wide range of signals: tapping the bucket,
pointing to the bucket, nodding to the bucket, or simply looking at the bucket.
The results showed that domestic dogs were better than chimpanzees, wolves, and human infants at this task, and even young
puppies with limited exposure to humans performed well. Dr. Stanley
Coren, an expert on dog
psychology, states that these results demonstrated the social cognition of dogs
can exceed that of even our closest genetic relatives, and that this capacity is
a recent genetic acquisition which distinguishes the dog from its ancestor, the
wolf. Studies have also investigated whether dogs engaged in partnered play
change their behavior depending on the attention-
Dr. Coren has also argued that dogs demonstrate a sophisticated theory of mind by engaging in deception, which he supports with a number of anecdotes, including one example where a dog hid a stolen treat by sitting on it until the rightful owner of the treat left the room. Although this could have been accidental, Coren suggests that the thief understood that the treat's owner would be unable to find the treat if it were out of view. Together, the empirical data and anecdotal evidence points to dogs possessing at least a limited form of theory of mind.
Behavior
Main article: Dog behavior
Further information: Category:Dog training and behavior
Although dogs have been the subject of a great deal of behaviorist psychology (e.g. Pavlov's dog), they do not enter the world with a psychological "blank slate". Rather, dog behavior is affected by genetic factors as well as environmental factors. Domestic dogs exhibit a number of behaviors and predispositions that were inherited from wolves. The Gray Wolf is a social animal that has evolved a sophisticated means of communication and social structure. The domestic dog has inherited some of these predispositions, but many of the salient characteristics in dog behavior have been largely shaped by selective breeding by humans. Thus some of these characteristics, such as the dog's highly developed social cognition, are found only in primitive forms in grey wolves.
Differences from wolves
Physical characteristics
Compared to equally sized wolves, dogs tend to have 20% smaller skulls, 30% smaller brains, as well as proportionately smaller teeth than other canid species. Dogs require fewer calories to function than wolves. Their diet of human refuse in antiquity made the large brains and jaw muscles needed for hunting unnecessary. It is thought by certain experts that the dog's limp ears are a result of atrophy of the jaw muscles. The skin of domestic dogs tends to be thicker than that of wolves, with some Inuit tribes favoring the former for use as clothing due to its greater resistance to wear and tear in harsh weather. The paws of a dog are half the size of those of a wolf, and their tails tend to curl upwards, another trait not found in wolves.
Behavior
Dogs tend to be poorer than wolves at observational learning, being more responsive to instrumental conditioning. Feral dogs show little of the complex social structure or dominance hierarchy present in wolf packs. For dogs, other members of their kind are of no help in locating food items, and are more like competitors. Feral dogs are primarily scavengers, with studies showing that unlike their wild cousins, they are poor ungulate hunters, having little impact on wildlife populations where they are sympatric. However, feral dogs have been reported to be effective hunters of reptiles in the Galápagos Islands, and free ranging pet dogs are more prone to predatory behavior toward wild animals.
Despite common belief, domestic dogs can be monogamous. Breeding in feral packs can be, but does not have to be
restricted to a dominant alpha pair (despite common belief, such things also
occur in wolf packs). Male dogs are unusual among canids by the fact that they
mostly seem to play no role in raising their puppies, and do not kill the young
of other females to increase their own reproductive success. Some sources say
that dogs differ from wolves and most other large canid species by the fact that
they do not regurgitate food for their young, nor the young of other dogs in the
same territory. However, this difference was not observed in all domestic dogs.
Regurgitating of food by the females for the young as well as care for the young
by the males has been observed in domestic dogs, dingos as well as in other
feral or semi-
Trainability
Dogs display much greater tractability than tame wolves, and are generally much more responsive to coercive techniques involving fear, aversive stimuli, and force than wolves, which are most responsive toward positive conditioning and rewards. Unlike tame wolves, dogs tend to respond more to voice than hand signals. Although they are less difficult to control than wolves, they can be comparatively more difficult to teach than a motivated wolf.
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